This study was supported by KOSEF NLRL Program grant 2011-0028333, Basic Science Research Program through NRF Korea grants 2012R1A2A2A01005939, 20100023233, and Global Frontier RD Program on < Human-centered Interaction for Coexistence > through NRF Korea grant NRF-2011-0032035. Image data collection was supported by NIH grants U01AG024904, P30AG010129, K01AG030514 and the Dana Foundation.